Publications of BCECB lab group at GXU

For full texts or other information, please see our ResearchGate accounts.


Guan, Z., Elleason, M., Goodale, E., and Mammides, C. 2021. Global patterns and drivers of human settlements within protected areas. Environmental Research Letters. DOI: 10.1088/1748-9326/ac056

Sreekar, R., Si, X., Sam, K., Liu, J., Dayananda, S., Goodale, U. M., Kotagama, S. W., Goodale, E. 2021. Land-use and elevation interact to shape bird functional and phylogenetic diversity and structure: Implications for designing optimal agriculture landscapes. Journal of Applied Ecology. DOI: 10.1111/1365-2664.13927.

Su, T., He, C., Jiang, A., Xu, Z., Goodale, E. and Qiu, G. 2021. Passerine bird reproduction does not decline in a highly-contaminated mercury mining area. Environmental Pollution 286: 117440.

Li, C., Chen, X., Jiang, A., Lee, M.-B., Mammides, C., and Goodale, E. 2021. Socioeconomic determinants of crop diversity, and its effect on farmer income in Guangxi, southern China. Agriculture 2021 (11): 336.

Mammides, C., Goodale, E., Dayananda, S., Luo, K., and Chen, J. 2021. On the use of the acoustic evenness index to monitor biodiversity: A comment on “Rapid assessment of avian species richness and abundance using acoustic indices” by Bradfer-Lawrence et al. (2020) [Ecological Indicators, 115, 106400]. Ecological Indicators 126: 107626

Meaux, E., Diana Castillo-Díaz, D., Pradhan, N., Dayananda, S. K., Balboa, C., Goodale, E., Mammides, C., Primack, R. B., Goodale, U. M. 2021. Transparency about human diversity in transnational environmental NGOs. Biological Conservation 256: 109027.

Martini, F., Aluthwattha, S. T., Mammides, C., Armani, M., and Goodale, U. M. 2021. Plant apparency drives leaf herbivory in seedling communities across four subtropical forests. Oecologica 195: 575–587.

Meaux, E., Peabotuwage, I., Mammides, C., Malykhina, K., Quan, R.-C., Goodale, E. 2021. Behavioural variables influence contact call rate more than characteristics of the vegetation in a group-living passerine species. Behavioural Processes 185: 104345.

Dai, W., Goodale, E., He, R., Mammides, C., Liu, S., Zhou, L., Tang, S., Su, B., Lao, X., Jiang, A. 2021. An eco-compensation policy increases shorebird diversity during the non-farming period for aquaculture. Wetlands 41: 4.

Sreekar, R., Sam, K., Dayananda, S. K., Goodale, U. M., Kotagama, S. W., and Goodale, E. 2021. Endemicity and land-use type influence the abundance–range size relationship of birds on a tropical island. Journal of Animal Ecology 90: 460-470.



Liang, D., Yang, S., Pagani-Núñez, E., He, C., Liu, Y., Goodale, E., Liao, W., Hu, J. 2020. How to become a generalist species? Individual niche variation across habitat transformation gradients. Frontiers in Ecology and Evolution 8: 597450.

Elleason, M., Guan, Z., Deng, Y., Jiang, A., Goodale, E., and Mammides, C. 2020. Strictly protected areas are not necessarily more effective than areas in which multiple human uses are permitted. Ambio 50: 1058-1073.

Zhou, X., Mammides, C., Zhou, X., Hobson, K.A., Zhu, H., Wen, Y., Jiang, A., 2020. Food web structure in exotic eucalyptus plantations in Southern China: Stable isotope (δ13C, δ15N) analyses reveal the importance of understory and landscape-level planning. Global Ecology and Conservation 24: e01259.

Bawa, K.S., Goodale, E., Mtemi, W., Chen, Y.-F., Barthakur, R., Goodale, U.M., Liu, J., Jiang, A., Mammides, C., Meegaskumbura, M., Pandit, M.K., Cao, K.-F., 2020. China and India: Toward a sustainable world. Science 369: 515.

Viswanathan, P., Mammides, C., Roy, P., Sharma, M. V., 2020. Flower visitors in agricultural farms of Nilgiri Biosphere Reserve: Do forests act as pollinator reservoirs? Journal of Apicultural Research 59: 978–987.

Zhang, Q., Holyoak, M., Goodale, E., Liu, Z., Shen, Y., Liu. J., Zhang, M., Dong, A., Zou, F. 2020. Trait-environment relationships differ between mixed-species flocking and non-flocking bird assemblages. Ecology 101: e03124.

Sreekar, R., Koh, L. P., Mammides, C., Corlett, R. T., Dayananda, S., Goodale, U. M., Kotagama, S. W., Goodale, E., 2020. Mechanisms assembling bird communities in the Western Ghats–Sri Lanka biodiversity hotspot are scale dependent: Roles of land-use, climate and distance. Oecologia 193: 801–809

Mammides, C., 2020. A global assessment of the human pressure on the world’s lakes. Global Environmental Change 63: 102084.

Mammides, C., 2020. A global analysis of the drivers of human pressure within protected areas at the national level. Sustainability Science 15, 1223–1232.

Anderson, E., Mammides, C., 2020. Changes in land-cover within high nature value farmlands inside and outside Natura 2000 sites in Europe: A preliminary assessment. Ambio 49: 1958–1971.

Jiang, D., Sieving, K. E., Meaux, E. and Goodale, E. 2020. Seasonal changes in mixed-species bird flocks and anti-predator information. Ecology and Evolution 10: 5368-5382.

Goodale, E., Sridhar, H., Sieving, K., Bangal, P., Colorado, G., Farine, D., Heymann, E. W., Jones, H., Krams, I., Martínez, A. E., Montaño-Centellas, F., Muñoz, J., Srinivasan, U., Theo, A., Shanker, K. 2020. Mixed company: A framework for understanding the composition and organization of mixed-species animal groups. Biological Reviews 95: 889-910.

Krams, I. A., Luoto, S., Krama, T., Krams, R., Sieving, K., Trakimash, G., Elfertsi, D., Rantalaj, M. J. and Goodale, E. 2020. Egalitarian mixed-species bird groups enhance winter survival of subordinate group members but only in high-quality forest. Scientific Reports 10: 4005.

Mammides, C., Kirkos, G., 2020. An analysis of the European Union’s conservation funding allocation by habitat and country. Environmental Conservation 47: 123–129.

Zhang, J.-Q., Mammides, C., Corlett, R.T., 2020. Reasons for the survival of tropical forest fragments in Xishuangbanna, Southwest China. Forests 11: 159.

Mammides, C. 2020. Evidence from eleven countries in four continents suggests that protected areas are not associated with higher poverty rates. Biological Conservation 241:108353.

Su, T., Lin, X., Huang, Q., Jiang, D., Zhang, C., Zhang, X., Dingle, C., Goodale, E., Que, P., Wang, R., and Liu, Y. 2020. Concentrations of mercury in sedentary and migratory Charadrius plovers distributed widely across China. Environmental Science and Pollution Research 27: 4236-4245.

He, C., Su, T., Liu, S., Jiang, A., Goodale, E., and Qiu, G. 2020. Heavy metal, arsenic and selenium concentrations in bird feathers from a region in southern China impacted by intensive mining of non-ferrous metals. Environmental Toxicology and Chemistry 39: 371-380.

Jiang, D., Goodale, E., Yang, G., Yu, L., Jiang, A., and Luo, X. 2020. Modeling the distribution of Nonggang Babbler (Stachyris nonggangensis), a threatened bird of limestone forests of the Sino-Vietnam border, and implications for its conservation. Bird Conservation International 30: 75-86.

Anderson, E. and Mammides, C. 2020. The role of protected areas in mitigating human impact in the world's last wilderness areas. Ambio 49: 434–441.



Lee, M.-B., Peabotuwage, I., Gu, H., Zhou, W., and Goodale, E. 2019. Factors affecting avian species richness and occupancy in a tropical city in southern China: Importance of human disturbance and open green space. Basic and Applied Ecology 39: 48-56.

Li, H., Goodale, E., and Quan, R. 2019. Nest predation on an abundant generalist bird in tropical China. Wilson Journal of Ornithology 131: 514-523.

Zhou, L., Peabotuwage, I., Gu, H., Jiang, D., Jiang, A., Zhang, M., Quan, R., and Goodale, E. 2019. The response of mixed-species flocks to anthropogenic disturbance and elevational variation in southwest China. Condor 121: duz028

Goodale, E., Ruxton, G. D., and Beauchamp, G. 2019. Predatory eavesdropping in a mixed-species environment: How prey species may use grouping, confusion, and the cocktail party problem to reduce predator detection. Frontiers in Ecology and Evolution 7:141.

Pagani-Núñez, E., Liang, D., He, C., Zhou, X., Luo, X., Liu, Y., and Goodale, E. 2019. Niches in the Anthropocene: Passerine assemblages show niche expansion from natural to urban habitats. Ecography 42: 1360-1369.

Mammides, C.  2019. European Union's conservation efforts are taxonomically biased. Biodiversity and Conservation 28: 1291-1296.

Priyadarshana, T.S., Atthanagoda, A.G., Wijewardhane, I.H., Siriweera, K.S., Aberathna, N., Kumar, P. 2019. Pteroceras dalaputtuwa (Orchidaceae: Epidendroideae: Vandeae: Aeridinae), a new species from Sri Lanka and re-collection of Pteroceras viridiflorum after 150 years. Phytotaxa 399: 65-76.

Peabotuwage, I., Goodale, U. M., and Goodale E. 2019. Is the keystone role of figs maintained across a gradient of increasing human disturbance? Biotropica 51: 300-303.

Tunheim, O.H., Stokke, B.G., Wang, L.W., Yang, C.C., Jiang, A.W., Liang, W., Roskaft, E., and Fossoy, F. 2019. Development and behavior of Plaintive Cuckoo (Cacomantis merulinus) nestlings and their Common Tailorbird (Orthotomus sutorius) hosts. Avian Research 10:5.

Ramachandran, V. S., Mohandass, D., Campbell, M.J., Mammides, C., and Shao, S.C. 2019. Selective logging intensity alters the population stand structure of Cullenia-Mesua-Palaquium dominated tropical wet evergreen forest of the Western Ghats, south India. Applied Ecology and Environmental Research 17: 9591-9607.

Qiu, G., Abeysinghe, K. S., Yang, X., Xu, Z., Xu, X., Luo, K., and Goodale, E. 2019. Effects of selenium on mercury bioaccumulation in a terrestrial food chain from an abandoned mercury mining region. Bulletin of Environmental Contamination and Toxicology 102: 329-334.

Goodale, E. and Ruxton, G. D.  2019. Antipredator benefits from heterospecifics. In: Encyclopedia of Animal Behavior, second edition (Choe, J.C., editor). Volume 4, pp. 400-406. Academic Press.


Mammides, C. 2018. Do satellite-derived data on forest loss correlate with indices of small-scale logging measured in the field? African Journal of Ecology 56: 390-394

Giosa, E., Mammides, C., and Zotos, S.  The importance of artificial wetlands for birds: A case study from Cyprus. PLoS ONE 13: e0197286.

Yuan, B.D., Yan, Y.F., Cheng, Z.Y., and Jiang, A.W. 2018. Roosting habitat selection of Hume's Pheasant (Syrmaticus humiae) in a fragmented forest patch, northwestern Guangxi, southwestern China. Global Ecology and Conservation 16: e00457.

Priyadarshana, T.S., Wijewardhane, I.H., Peabotuwage, I., Jayasooriya, A., and Herath, B.E. 2018. A new species of Ceylonosticta Fraser, 1931 (Odonata: Zygoptera: Platystictidae) from Sri Lanka. International Journal of Odontology 21: 105-114.

Hai, B.T., Chen, J., McConkey, K.R. and Dayananda, S.K. 2018. Gibbons (Nomascus gabriellae) provide key seed dispersal for the Pacific walnut (Dracontomelon dao), in Asia's lowland tropical forest. Acta Oecologia 88: 71-79.

Liang, D., He, C.°, Luo, X., Liu, Y.*, Goodale, E., and Pagani-Núñez, E*. Breath rate of passerines across an urbanization gradient supports the pace-of-life hypothesis and suggests diet-mediated responses to handling stress. Ecology and Evolution. DOI: 10.1002/ece3.4460

Lee, M.-B.* and Goodale, E. 2018. Crop heterogeneity and non-crop habitat features can enhance avian diversity in a tropical agricultural landscape in southern China. Agriculture, Ecosystems and Environment 265: 254-263.

Mammides, C.*, Chen, J., Goodale, U. M., Kotagama, S. W., and Goodale, E. 2018. Measurement of species associations in mixed-species bird flocks across environmental and human disturbance gradients.  Ecosphere 9: e02324.

Zou, F.*, Jones, H., Colorado Z., G. J., Jiang, D. #, Lee, T.-M., Martínez, A., Sieving, K., Zhang, M., Zhang, Q., and Goodale, E.* 2018. The conservation implications of mixed-species flocking in terrestrial birds, a globally-distributed species interaction network. Biological Conservation 224: 267-276

Zhou, W., Lee, M.-B.*, and Goodale, E*. 2018. The relationship between the diversity of herbaceous plants and the extent and heterogeneity of croplands in noncrop vegetation in an agricultural landscape of south China. Global Ecology and Conservation 14: e000399.

Pagani-Núñez, E., Xia, X., Beauchamp, G., He, R., Husson, J. H. D., Liang, D., and Goodale, E. 2018. Are vocal characteristics related to leadership patterns in mixed-species bird flocks? Journal of Avian Biology. doi: 10.1111/jav.01674

Xing, S., Cheng, W., Nakamura, A., Tang, C. C., Pickett, E., Huang, S., Odell, E., Goodale, E., Goodale, U. M., and Bonebrake, T. 2018. Elevational clines in morphological traits of subtropical and tropical butterfly assemblages. Biological Journal of the Linnean Society 123: 506-517.



Priyadarshana, T. S., Wijewardhane, I. H., & Karunarathna, M. (2017). A note on the distribution of two highly threatened butterflies in Sri Lanka (Lepidoptera: Lycaenidae: Spindasis greeni and Rapala lankana), with a report on the range extension of S. greeni. Journal of Threatened Taxa, 9(11), 10971-10973. DOI: 10.11609/jott.3274.9.11.10971-10973. 

Jiang, A., Zhou, P., Jiang, D., He, C., Yu, M., and Zeng Q. 2017. Species composition, fauna and elevational gradient pattern of birds in Damingshan, southern central Guangxi. Chinese Journal of Zoology 52: 177-193.

Jenkins, H.M., Mammides, C., and Keane, A. 2017. Exploring differences in stakeholders' perceptions of illegal bird trapping in Cyprus. Journal of Ethnobiology and Ethnomedicine 13: 67.

Jiang, A., Jiang, D., Zhou, F., and Goodale, E. 2017. Nest-site selection and breeding ecology of Streaked Wren-Babbler (Napothera brevicaudata) in a tropical limestone forest of southern China. Avian Research 8: 28.

Dayananda, S. K., Mammides, C., Lee, M.-B., Liu, J.-J., Pasion, B. O., Sreekar, R., Yasuda, M., Quan, R.-C., Slik, J. W. F., Tomlinson, K. W. and Goodale, E. 2017. Topography and soil type are critical to understanding how bird and herpetofaunal communities persist in forest fragments of tropical China. Biological Conservation 215: 107-115.

Souter, N. J., Hughes, A. C., Savini, T., Rao, M., Goodale, E., Van Nice, A., Huang, N., Liu, J.-X., Hunt, M. P., O’Connor, D. A., Hueng-Lam, A. L., Gnuen, G., Sun, Y., and Silva, I.  2017. Building capacity for conservation in Southeast Asia: outcomes of the ATBC 2015 Asia-Pacific Chapter meeting conservation education symposium. Applied Environmental Education and Communication. doi: 10.1080/1533015X.2017.1322012

Mammides, C., Goodale, E., Dayananda, S. K., Kang, L. and Chen, J. 2017. Do acoustic indices correlate with bird diversity? Insights from two biodiverse areas in Yunnan Province, south China. Ecological Indicators 82: 470-477.

Mohandass, D., Campbell, M. J., Hughes, A. C., Mammides, C. and Davidar, P.  The effect of altitude, patch size and disturbance on species richness and density of lianas in montane forest patches. Acta Oecologia 83: 1-14.

Goodale, E. 2017. Who does conservation science and why that matters: A personal perspective. Current Conservation 11: 11-14.

Ding, Z. , Ji, F.,  Huang, Q. Wang, L., Jiang, A., Zhang, C., Feng, Y., Tian, Y., Huijian Hu, H., and Liang, W. 2017. Brood sex ratio in the Yellow‑bellied Prinia (Prinia flaviventris). Avian Research 8: 15.

Abeysinghe, K., Qiu, G., Goodale, E., Anderson, C. W. N., Bishop, K., Du, B., Evers, D. C., Goodale, M. W., Hintelmann, H., Li, P., Liu, S., Mammides, C., Quan, R.-C, Wang, J., Wu, P., Xu, X.-H., Yang, X.-D., Xiong, M. and Feng, X. 2017. Mercury flow through an Asian rice-based food web. Environmental Pollution 229: 219-228.

Gong, S., Shi, A., Jiang, A., Fong, J. J., Gaillard, D., and Wang, J. Disappearance of endangered turtles within China’s nature reserves. Current Biology 27: R170-R171.

He, R., Pagani-Núñez, E., Chevallier, C., and Barnett, C. R. A. To be so bold: Boldness is repeatable and related to within individual behavioural variability in North Island robins. Behavioral Processes 140: 144-149.

Pagani-Núñez, E., He, C., Wu, Y. W., Peabotuwage, I., and Goodale, E. 2017. Foraging in the tropics: Relationships between species’ abundances, niche asymmetries and body condition in an urban avian assemblage. Urban Ecosystems. DOI: 10.1007/s11252-017-0682-1

Sreekar, R., Corlett, R., Dayananda, S., Goodale, U. M., Kilpatrick, A., Kotagama, S. W., Koh, L. P., and Goodale, E. 2017. Horizontal and vertical species turnover in tropical birds in habitats with differing land use. Biology Letters 13: 20170186.

Priyadarshana, T. S., Wijewardhane, I. H., Atthanagoda, A. G., Arangala, N. S., Jayasooriya, A., and Kumar, O. 2017. Oberonia meegaskumburae (Orchidaceae: Epidendroideae: Malaxideae), a new species from Sri Lanka. Phytotaxa 302: 259-265.

Priyadarshana, T. S., and Wijewardhane, I. H. 2017. Lifecycle and fecula measurements of Cheritra freja (Lepidoptera: Lycaenidae), as relevant to the different larval stages. Journal of Threatened Taxa 9: 10011-10013.

Gu, H., Chen, J., Ewing, H., Liu, X., Zhao, J., and Goodale, E. 2017. Heterospecific attraction to the vocalizations of birds in mass fruiting trees. Behavioral Ecology and Sociobiology 71: 82.

Jiang, A., Jiang, D., Wen, Y. and Goodale. E. 2017. Nest predation on birds that nest in rock cavities in a tropical limestone forest of southern China. Global Ecology and Conservation 10: 154-158.

Jiang, A., Tan, L., and Feng, H. 2017.  Breeding observations of White-Eared Night-Herons (Gorsachius magnificus) in artificial forests of southern China. Waterbirds 40: 173-179.

Jiang, A., Yang, G., Pagani-Núñez, E., and Jiang, D. 2017. Ecology of two pittas (Pitta soror and Pitta nympha) in limestone forests of south China. Journal of Natural History. DOI:

Goodale, E., Beauchamp, G. and Ruxton, G. D. 2017. Mixed-species animal groups: Behavior, community ecology and conservation.  Academic Press (book). ISBN: 978-0-12-805355-3.

Abeysinghe, K., Yang, X.-D., Goodale, E., Anderson, C. W. N., Bishop, K., Cao, A., Feng, X., Liu, S., Mammides, C., Meng, B., Quan, R.-C, Sun, J., and Qiu, G. 2017. Total and methyl mercury concentrations over a gradient of contamination in earthworms living in rice paddy soil. Environmental Toxicology and Chemistry 36: 1202-1210.



McCollum, K. R., Conway, A. L., Lee, M.-P., and Carroll, J. P.. 2016. Occupancy and demographics of red river hog Potamochoerus porcus on Tiwai Island, Sierra Leone. African Journal of Ecology 55: 47-55

Perera, M. P., Kotagama, S. W., Goodale, E., and Kathriarachchi, H. S. 2016.  What happens when the nuclear species is absent?: Observations of mixed-species bird flocks in the Hiyare Forest Reserve in Sri Lanka. Forktail 32: 96-97.

Jiang, A., Jiang, D., Goodale, E., Zhou, F., Wen, H. 2016. Olive-backed Sunbird Cinnyris jugularis assisting Crested Bunting Melophus lathami at the nest: substantiated evidence for interspecific feeding, Guangxi, south-west China. Forktail 32: 93-96

Yang, C., Huang, Q., Wang, L., Jiang, A., Stokke, B. G., Fossøy, F., Tunheim, O. H., Røskaft, E., Liang, W., Møller, A. P. 2016.  Plaintive cuckoos do not select tailorbird hosts that match the phenotypes of their own eggs. Behavioral Ecology 27: 275-279.

Priyadarshana, T. S., Wijewardhana, I. H., and Herath, B. E. 2016. Three new species of the genus Ceylonosticta Fraser, 1931 (Odonata: Zygoptera: Platystictidae) from Sri Lanka and the rediscovery of Ceylonosticta subtropica (Fraser, 1933). International Journal of Odonatology. DOI: 10.1080/13887890.2016.1257443

Duguid, M., Morrell, E. H., Goodale, E., and Ashton, M. S.  2016. Changes in breeding bird abundance and species composition over a 20 year chronosequence following shelterwood harvests in oak-hardwood forests. Forest Ecology and Management 376: 221-230.

Pagani-Nuñéz, E., Barnett, C. A., Gu, H. and Goodale, E. 2016. The need for new categorizations of dietary specialism incorporating spatio-temporal variability of individual diet specialization. Journal of Zoology 300: 1-7.

Pagani-Nuñéz, E., Chao, H., Li, B., Li, M., He, R., Jiang, A., and Goodale, E. 2016. The breeding ecology of the barn swallow Hirundo rustica gutturalis in South China. Journal of Tropical Ecology 32: 260-263.  

Dayananda, S. K., Goodale, E., Liu, J.-J., Mammides, C., Pasion, B., Quan, R.-C., Slik, J. W. F., Sreekar, R., Tomlinson, K. W., and Yasuda, M. 2016. Effects of forest fragmentation on nocturnal birds: A case study from Xishuangbanna, China. Zoological Research 37: 151-158.

Mammides, C., Goodale, U. M., Corlett, R. T., Chen, J., Bawa, K. S., Hariya, H., Jarrad, F., Primack, R. B., Ewing, H., Xia, X. and Goodale, E. 2016. Increasing geographic diversity in the international conservation literature: A stalled process? Biological Conservation 198: 78-83.

Peiró, I. and Pagani-Núñez, E. 2016. Plumage coloration variability of male Bluethroats (Luscinia svecica cyanecula) wintering in SE Iberia. Ornis Fennica 93: 111-120.

Sreekar, R., Huang, G., Yasuda, M., Quan, R.-C., Goodale, E., Corlett, R. T. and Tomlinson, K. W. 2016. Can rubber monocultures be made more hospitable to birds? The effects of forests, roads and mistletoes. Scientific Reports 6: 21822.

Mammides, C., Kounnamas, C., Goodale, E., and Kadis, K. 2016. Do unpaved, low-traffic roads affect bird communities? Acta Oecologica 71: 14-21.



Kassinis, N. and Mammides, C. 2015. Winter bird surveys in Cyprus, 2007–2016. Analysis of the population trends. Bird Census News 29: 37-45.

Quan, R.-C., Li, H., Wang, B. and Goodale, E. 2015. The relationship between defecation and feeding in nestling birds: Observational and experimental evidence. Frontiers in Zoology 12:21.

Goodale, E., Ding, P., Liu, X., Martínez, A., Walters, M. & Robinson, S. K. 2015. The structure of mixed-species bird flocks, and their response to anthropogenic disturbance, with special reference to East Asia. Avian Research 6:14.


2015 articles with XTBG address

Yang, H., Harrison, R., Yi, Z.-F., Goodale, E., Zhao, M.-X., and Xu, J.-C. 2015. Changing perceptions of forest value and attitudes toward management of a recently established nature reserve: a case study in southwest China. Forests 6: 3136-3164.

Mammides, C., Chen, J., Goodale, U. M., Kotagama, S. W., Sidhu, S. & Goodale, E. 2015. Does mixed-species flocking influence how birds respond to a gradient of land-use intensity? Proceedings of the Royal Society B, Biological Sciences 282: 20151118.

Sreekar, R., Goodale, E., and Harrison, R. D. 2015. The use of flight initiation distance as index of hunting pressure: a case study in Xishuangbanna, southwest China. Tropical Conservation Science 8: 505-512.

Sreekar, R., Srinivasan, U., Mammides, C., Chen, J., Goodale, U.M., Kotagama, S. W., Sidhu, S. and Goodale, E. 2015. The effect of land-use on the diversity and mass-abundance relationships of understory avian insectivores in Sri Lanka and southern India. Scientific Reports 5: 11569.

Gu, H., Goodale, E. & Chen, J. 2015. Emerging directions in the study of the ecology and evolution of plant-animal mutualistic networks: a review. Zoological Research 36: 65-71.

Gu, H., Goodale, E. & Chen, J. 2015. Does the role that frugivorous bird species play in seed dispersal networks influence their rate of evolutionary divergence? Global Ecology and Conservation 3: 121-128.

Sreekar, R., Huang, G., Zhao, J.- B., Pasion, B. O., Yasuda, M., Zhang, K., Peabotuwage, I.,  Wang, X.,  Quan, R.- C., Slik, F., Corlett, R. T., Goodale, E. & Harrison, R. D. 2015. The use of species-area relationships to partition the effects of hunting and deforestation on bird extirpations in a fragmented landscape. Diversity and Distributions 21: 441-450.


Publications of Prof Goodale 2014 and before

Duan, Q., Goodale, E. & Quan R.-C. 2014. Bird fruit preferences reflect the frequency of fruit colours in tropical Asia. Scientific Reports 4: 5627.

Zhang, W., Goodale, E. & Chen, J. 2014. How contact with nature affects children’s biophilia, biophobia and conservation attitude in China. Biological Conservation 177: 109-116.

Goodale, E., Kim, E., Nabors, A., Henrichon, S.  & Nieh, J. C. 2014. The innate responses of bumble bees to flower patterns: separating the nectar guide from the nectary changes bee movements and search time. Naturwissenschaften 101: 523-526.

Zhang K., Woan, T. S., Jie, L., Goodale, E., Kitajima, K. & Harrison, R. D. 2014. Shifting baselines on a tropical forest frontier: Extirpations drive declines in local ecological knowledge. PLOS ONE 9: e86598.

Goodale, E., Ratnayake, C. P. & Kotagama, S. W. 2014. Vocal mimicry of alarm-associated sounds by a drongo elicits flee and mobbing responses from other species that participate in mixed-species bird flocks. Ethology 120: 266-274.

Goodale, E., Ratnayake, C. P. & Kotagama, S. W. 2014. The frequency of vocal mimicry varies due to proximity to nest and nesting stage in a passerine bird. Behaviour 151: 73-88.

Goodale, E., Kotagama, S. W., Raman, T. R. S., Sidhu, S., Goodale, U. M., Parker, S. & Chen, J. 2014. The response of birds and mixed-species bird flocks to human-modified landscapes in Sri Lanka and southern India. Forest Ecology and Management 329: 384-392.

Jayarathna, A., Kotagama, S. W. & Goodale, E. 2013. The seasonality of mixed-species flocking in a Sri Lankan rainforest, in relation to the breeding of the nuclear species, the Orange-billed Babbler Turdoides rufescens. Forktail 29: 138-139.

Goodale, E. & Nieh, J. C. 2012. Public use of olfactory information associated with predation in two species of social bees. Animal Behaviour 84: 919-924.

Sridhar, H., Srinivasan, U., Askins, R. A., Canales Delgadillo, J. C., Chen, C.-C., Ewert, D. N., Gale, G. A., Goodale, E., Gram, W. K., Hart, P. J.,  Hobson, K. A., Hutto, R. L., Kotagama, S. W., Knowlton, J., Lee, T. M., Munn, C. A., Nimnuan, S., Nizam, B. Z., Péron, G., Robin, V. V., Rodewald, A. D., Rodewald, P. G., Thomson, R. L., Trivedi, P., Van Wilgenburg, S. L. & Shanker, K. 2012. Positive relationships between association strength and phenotypic similarity characterize the assembly of mixed-species bird flocks globally. American Naturalist 180: 777-790.

Goodale, E., Goodale, U. & Mana, R. 2012. The role of toxic pitohuis in mixed-species flocks of lowland forest in Papua New Guinea. Emu – Austral Ornithology 112: 9-16.

Beauchamp, G. & Goodale, E. 2011. Plumage mimicry in avian mixed-species flocks: more or less than meet the eye? Auk 128: 487-496.

Sidhu, S., Raman, T. R. S. & Goodale, E. 2010. Effects of plantations and home-gardens on tropical forest bird communities and mixed-species bird flocks in the southern Western Ghats. Journal of the Bombay Natural History Society 107: 91-108

Goodale, E. & Ruxton, G. D.  2010. Antipredator benefits from heterospecifics. In: Encyclopedia of Animal Behavior (Breed, M and Moore, J., editors). Academic Press, Oxford.

Goodale, E. & Podos, J. 2010. Persistence of song types in Darwin’s finches, Geospiza fortis, over four decades. Biology Letters 6: 589-592.

Goodale, E., Beauchamp, G., Magrath, R. D., Nieh, J. C. & Ruxton, G. D. 2010. Interspecific information transfer influences animal community structure. Trends in Ecology and Evolution 25: 354-361.

Satischandra, S. H. K., Kodituwakku, P., Kotagama, S. W. & Goodale, E.  2010. Assessing “false” alarm calls by a drongo (Dicrurus paradiseus) in mixed-species bird flocks. Behavioral Ecology 21: 396-403.

Goodale, E & Beauchamp, G.  2010. The relationship between leadership and gregariousness in mixed-species bird flocks. Journal of Avian Biology 41: 99-103.

Ratnayake, C. P., Goodale, E. & Kotagama, S. W.  2010. Two sympatric species of passerine birds imitate the same raptor calls in alarm contexts. Naturwissenschaften 97: 103-108.

Goodale, E., Nizam, B. Z., Robin, V. V., Sridhar, H., Trivedi, P., Kotagama, S. W., Padmalal, U. K. G. K., Perera, R., Pramod, P. & Vijayan, L. 2009. Regional variation in structure and composition of mixed-species bird flocks of the Western Ghats and Sri Lanka: a review. Current Science 97: 648-663.

Goodale, E., Lalbhai, P., Goodale, U. M., & Ashton, P. M. S. 2009. The effect of shelterwood cuts and crown thinnings on the abundance and distribution of birds in a southern New England forest. Forest Ecology and Management 258: 314-322.

Goodale, E. & Kotagama, S.W. 2008.  Responses to conspecific and heterospecific alarm calls in mixed-species bird flocks of a Sri Lanka rainforest. Behavioral Ecology 19: 887-894.

Goodale, E. & Kotagama, S.W.  2007.  Some observations on the geographic variation of mixed-species bird flocks in Sri Lanka. Journal of the Bombay Natural History Society 104: 96-98.

Joachim, D. & Goodale, E.  2007. On the use of cellular telephony for audio interaction with animals. Biology Letters 3: 603-606.

Sathischandra, S.H.K., Kudavidanage, E.P., Kotagama, S.W. & Goodale, E.  2007. The benefits of joining mixed-species flocks for a sentinel nuclear species, the Greater Racket-tailed Drongo Dicrurus paradiseus. Forktail 23: 145-148.

Goodale, E. & Kotagama, S.W.  2006. Vocal mimicry by a passerine bird attracts other species involved in mixed-species flocks. Animal Behaviour 72: 471–477.

Goodale, E. & Kotagama, S.W. 2006. Context-dependent mimicry by a passerine bird. Proceedings of the Royal Society B, Biological Sciences 273: 875–880.

Goodale, E. & Kotagama, S.W. 2005. Alarm calling in Sri Lankan mixed-species bird flocks. Auk 122: 108–120.

Goodale, E. & Kotagama, S.W. 2005.  Testing the roles of species in mixed-species bird flocks of a Sri Lankan rainforest. Journal of Tropical Ecology 21: 669–676.

Jayarathna, A., Caldera, P. & Goodale, E.  2005.  Observations on the nesting and parental behaviour of the Ashy-headed Laughing Thrush (Garrulax cinereifrons). Forktail 21: 163­–164.

Kotagama, S.W. & Goodale, E.  2004.  The composition and spatial organization of mixed-species flocks in a Sri Lankan rainforest. Forktail 20: 63–70.

Kroodsma, D.E., Byers, B.E., Goodale, E., Johnson, S. & Liu, W.   2001. Pseudoreplication in playback experiments, revisited a decade later. Animal Behaviour 61: 1029­–1033.